@article{MTMT:30646377,
	title = {Karst dolines provide diverse microhabitats for different functional groups in multiple phyla},
	url = {https://m2.mtmt.hu/api/publication/30646377},
	author = {Bátori, Zoltán and Vojtkó, András and Maák, István Elek and Lőrinczi, Gábor and Farkas, T. and Kántor, N. and Tanács, Eszter and Kiss, Péter János and Juhász, Orsolya and Módra, Gábor and Tölgyesi, Csaba and Erdős, László and Aguilon, Dianne Joy and Keppel, G.},
	doi = {10.1038/s41598-019-43603-x},
	journal-iso = {SCI REP},
	journal = {SCIENTIFIC REPORTS},
	volume = {9},
	unique-id = {30646377},
	issn = {2045-2322},
	abstract = {Fine-scale topographic complexity creates important microclimates that can facilitate species to grow outside their main distributional range and increase biodiversity locally. Enclosed depressions in karst landscapes (‘dolines’) are topographically complex environments which produce microclimates that are drier and warmer (equator-facing slopes) and cooler and moister (pole-facing slopes and depression bottoms) than the surrounding climate. We show that the distribution patterns of functional groups for organisms in two different phyla, Arthropoda (ants) and Tracheophyta (vascular plants), mirror this variation of microclimate. We found that north-facing slopes and bottoms of solution dolines in northern Hungary provided key habitats for ant and plant species associated with cooler and/or moister conditions. Contrarily, south-facing slopes of dolines provided key habitats for species associated with warmer and/or drier conditions. Species occurring on the surrounding plateau were associated with intermediate conditions. We conclude that karst dolines provide a diversity of microclimatic habitats that may facilitate the persistence of taxa with diverse environmental preferences, indicating these dolines to be potential safe havens for multiple phyla under local and global climate oscillations.},
	year = {2019},
	eissn = {2045-2322},
	orcid-numbers = {Bátori, Zoltán/0000-0001-9915-5309; Maák, István Elek/0000-0002-0999-4916; Lőrinczi, Gábor/0000-0001-5384-7246; Tanács, Eszter/0000-0003-1953-9340; Tölgyesi, Csaba/0000-0002-0770-2107; Erdős, László/0000-0002-6750-0961}
}

@article{MTMT:3157928,
	title = {Large- and small-scale environmental factors drive distributions of cool-adapted plants in karstic microrefugia},
	url = {https://m2.mtmt.hu/api/publication/3157928},
	author = {Bátori, Zoltán and Vojtkó, András and Farkas, T and Szabó, A and Havadtői, K and E-Vojtkó, Anna and Tölgyesi, Csaba and Cseh, Viktória and Erdős, László and Maák, István Elek and Keppel, G},
	doi = {10.1093/aob/mcw233},
	journal-iso = {ANN BOT-LONDON},
	journal = {ANNALS OF BOTANY},
	volume = {119},
	unique-id = {3157928},
	issn = {0305-7364},
	abstract = {Background and aims Dolines are small-to large-sized bowl-shaped depressions of karst surfaces. They may constitute important microrefugia, as thermal inversion often maintains cooler conditions within them. This study aimed to identify the effects of large-(macroclimate) and small-scale (slope aspect and vegetation type) environmental factors on cool-adapted plants in karst dolines of East-Central Europe. We also evaluated the potential of these dolines to be microrefugia that mitigate the effects of climate change on cool-adapted plants in both forest and grassland ecosystems. Methods We compared surveys of plant species composition that were made between 2007 and 2015 in 21 dolines distributed across four mountain ranges (sites) in Hungary and Romania. We examined the effects of environmental factors on the distribution and number of cool-adapted plants on three scales: (1) regional (all sites); (2) within sites and; (3) within dolines. Generalized linear models and non-parametric tests were used for the analyses. Key Results Macroclimate, vegetation type and aspect were all significant predictors of the diversity of cooladapted plants. More cool-adapted plants were recorded in the coolest site, with only few found in the warmest site. At the warmest site, the distribution of cool-adapted plants was restricted to the deepest parts of dolines. Within sites of intermediate temperature and humidity, the effect of vegetation type and aspect on the diversity of cool-adapted plants was often significant, with more taxa being found in grasslands (versus forests) and on north-facing slopes (versus south-facing slopes). Conclusions There is large variation in the number and spatial distribution of cool-adapted plants in karst dolines, which is related to large- and small-scale environmental factors. Both macro-and microrefugia are therefore likely to play important roles in facilitating the persistence of cool-adapted plants under global warming.},
	keywords = {VEGETATION; Central-Europe; BALKAN PENINSULA; East-central Europe; environmental gradient; Refugia; Refugia; COMPARATIVE PHYLOGEOGRAPHY; CLIMATE-CHANGE IMPACTS; RANGE SHIFTS; Capacity of refugia; high-mountain plants; karst dolines; migration processes; relicts; MOUNTAINS SOUTH HUNGARY; ALPINE PLANTS},
	year = {2017},
	eissn = {1095-8290},
	pages = {301-309},
	orcid-numbers = {Bátori, Zoltán/0000-0001-9915-5309; E-Vojtkó, Anna/0000-0001-6370-680X; Tölgyesi, Csaba/0000-0002-0770-2107; Erdős, László/0000-0002-6750-0961; Maák, István Elek/0000-0002-0999-4916}
}

@article{MTMT:3159453,
	title = {Stand structural drivers of microclimate in mature temperate mixed forests},
	url = {https://m2.mtmt.hu/api/publication/3159453},
	author = {Kovács, Bence and Tinya, Flóra and Ódor, Péter},
	doi = {10.1016/j.agrformet.2016.11.268},
	journal-iso = {AGR FOREST METEOROL},
	journal = {AGRICULTURAL AND FOREST METEOROLOGY},
	volume = {234},
	unique-id = {3159453},
	issn = {0168-1923},
	year = {2017},
	eissn = {1873-2240},
	pages = {11-21},
	orcid-numbers = {Kovács, Bence/0000-0002-8045-8489; Tinya, Flóra/0000-0002-4271-9676; Ódor, Péter/0000-0003-1729-8897}
}

@article{MTMT:2725970,
	title = {A comparison of the vegetation of forested and non-forested solution dolines in Hungary: a preliminary study},
	url = {https://m2.mtmt.hu/api/publication/2725970},
	author = {Bátori, Zoltán and Farkas, T. and Erdős, László and Tölgyesi, Csaba and Körmöczi, László and Vojtkó, András},
	doi = {10.2478/s11756-014-0430-4},
	journal-iso = {BIOLOGIA},
	journal = {BIOLOGIA (BRATISLAVA)},
	volume = {69},
	unique-id = {2725970},
	issn = {0006-3088},
	abstract = {The present study compares the vegetation characteristics of two large forested and one large non-forested solution dolines in Hungary. We investigated the species composition and vegetation pattern along north to south transects (across the doline bottoms) and compared the richness of different species groups (dry and wet groups) on the doline slopes. We applied linear regression models for each slope to explore the effects of topography on species richness, and Detrended Correspondence Analysis (DCA) to detect the major gradients of floristic variation within each site. We found that the vegetation changed significantly along all transects; and, regardless of the vegetation cover, the doline bottoms contained several cool-adapted species. Variations within the two species groups were more pronounced on the south-facing slopes. The changes were similar in the forested dolines, indicating the role of forest cover in maintaining many cool-adapted species on the north-facing slopes as well. However, the number of cool-adapted species increased significantly along both slopes of the non-forested doline from the upper edge to the bottom. Contrary to our expectations, the species turnover along the slopes of the non-forested doline was lower than that along the slopes of the forested ones. We conclude that both the forested and non-forested dolines serve as refuges for many plant species adapted to different environmental conditions. Apart from providing an understanding of population patterns along environmental gradients, our results may also contribute to our understanding of an even more fundamental question for a future research agenda: the probable effects of climate change on vegetation characteristics in climatic islands with environmental conditions substantially different from the surrounding areas.},
	keywords = {SOIL; climate; climate change; Mediterranean Region; refugium; Microclimate; Relict species; MOUNTAINS SOUTH HUNGARY; karst surface; KARSTIC SINKHOLES; SLOPE ASPECT},
	year = {2014},
	eissn = {1336-9563},
	pages = {1339-1348},
	orcid-numbers = {Bátori, Zoltán/0000-0001-9915-5309; Erdős, László/0000-0002-6750-0961; Tölgyesi, Csaba/0000-0002-0770-2107; Körmöczi, László/0000-0003-1027-3822}
}

@article{MTMT:2453551,
	title = {The conservation value of karst dolines for vascular plants in woodland habitats of Hungary: refugia and climate change},
	url = {https://m2.mtmt.hu/api/publication/2453551},
	author = {Bátori, Zoltán and Csiky, János and Farkas, Tünde and E-Vojtkó, Anna and Erdős, László and Kovács, Dániel and Wirth, Tamás and Körmöczi, László and Vojtkó, András},
	doi = {10.5038/1827-806X.43.1.2},
	journal-iso = {INT J SPELEOL},
	journal = {INTERNATIONAL JOURNAL OF SPELEOLOGY},
	volume = {43},
	unique-id = {2453551},
	issn = {0392-6672},
	year = {2014},
	eissn = {1827-806X},
	pages = {15-26},
	orcid-numbers = {Bátori, Zoltán/0000-0001-9915-5309; E-Vojtkó, Anna/0000-0001-6370-680X; Erdős, László/0000-0002-6750-0961; Körmöczi, László/0000-0003-1027-3822}
}

@article{MTMT:2485324,
	title = {Microhabitat associations of land snails in forested dolinas: implications for coarse filter conservation},
	url = {https://m2.mtmt.hu/api/publication/2485324},
	author = {Kemencei, Zita and Farkas, Roland Tibor and Páll-Gergely, B and Vilisics, Ferenc and Nagy, Antal and Hornung, Erzsébet and Sólymos, Péter},
	doi = {10.1556/comec.15.2014.2.6},
	journal-iso = {COMMUNITY ECOL},
	journal = {COMMUNITY ECOLOGY},
	volume = {15},
	unique-id = {2485324},
	issn = {1585-8553},
	year = {2014},
	eissn = {1588-2756},
	pages = {180-186},
	orcid-numbers = {Sólymos, Péter/0000-0001-7337-1740}
}

@article{MTMT:2041669,
	title = {Importance of karst sinkholes in preserving relict, mountain, and wetwoodland plant species under sub-Mediterranean climate: A case study from southern Hungary},
	url = {https://m2.mtmt.hu/api/publication/2041669},
	author = {Bátori, Zoltán and Körmöczi, László and Erdős, László and Zalatnai, Márta and Csiky, János},
	doi = {10.4311/2011LSC0216},
	journal-iso = {J CAVE KARST STUD},
	journal = {JOURNAL OF CAVE AND KARST STUDIES},
	volume = {74},
	unique-id = {2041669},
	issn = {1090-6924},
	year = {2012},
	eissn = {2331-3714},
	pages = {127-134},
	orcid-numbers = {Bátori, Zoltán/0000-0001-9915-5309; Körmöczi, László/0000-0003-1027-3822; Erdős, László/0000-0002-6750-0961}
}

@article{MTMT:1744742,
	title = {Seminatural grassland management by mowing of calamagrostis epigejos in Hungary},
	url = {https://m2.mtmt.hu/api/publication/1744742},
	author = {Házi, Judit and Bartha, Sándor and Szentes, Szilárd and Wichmann, Barnabás and Penksza, Károly},
	doi = {10.1080/11263504.2011.601339},
	journal-iso = {PLANT BIOSYST},
	journal = {PLANT BIOSYSTEMS},
	volume = {145},
	unique-id = {1744742},
	issn = {1126-3504},
	abstract = {Regeneration of seminatural grasslands are often threatened by the invasion of Calamagrostis epigejos, which can slow down or arrest secondary succession. Here we report the results of a 9-year mowing experiment designed to suppress the spread of C. epigejos in mid-successional grasslands in Hungary. The experimental design consisted of 16 permanent plots of 3 × 3 m. Half of the plots were mowed twice a year (in June and September), the other half was left as control. Vegetation was sampled in 2 × 2 m quadrates before mowing in each year between 2001 and 2009. The effects of mowing were tested using repeated-measure analyses of variance (ANOVA) and Tukey HSD for post hoc tests. Significant decrease of C. epigejos appeared after 2 years of mowing. Species richness increased after 4 years, while diversity after 8 years. By this time the target native species Brachypodium pinnatum become dominant. Similar trends appeared in the control plots during spontaneous succession but at much slower rates. Our results suggest that C. epigejos disappears spontaneously in secondary grassland succession after ca. 40-50 years. However, mowing twice a year can speed up this process by opening a "colonization window" to the valuable target species. For successful control, mowing should be maintained for approximately 8 years. © 2011 Società Botanica Italiana.},
	keywords = {RESTORATION; Biodiversity; Spontaneous succession; Secondary meadow steppe; Forest-steppe zone; Abandoned vineyards},
	year = {2011},
	eissn = {1724-5575},
	pages = {699-707},
	orcid-numbers = {Bartha, Sándor/0000-0001-6331-7521}
}

@book{MTMT:1395109,
	title = {Magyarország kistájainak katasztere. Második, átdolgozott és bővített kiadás},
	url = {https://m2.mtmt.hu/api/publication/1395109},
	isbn = {9789639545298},
	author = {Becse, A and Mezősi, Gábor and Ádám, László and Juhász, Ágoston and Marosi, S and Somogyi, Sándor and Szilárd, J and Ambrózy, P and Konkolyné, Bihari Z and Király, Botond Gergely and Molnár, Zsolt and Bölöni, János and Csiky, János and Vojtkó, András and Rajkai, Kálmán László and Tóth, Gergely and Tiner, Tibor and Dövényi, Zoltán and Michalkó, Gábor and Keresztesi, Zoltán},
	editor = {Dövényi, Zoltán},
	publisher = {Magyar Tudományos Akadémia Földrajztudományi Kutatóintézet; MTA FKI},
	unique-id = {1395109},
	year = {2010},
	orcid-numbers = {Király, Botond Gergely/0000-0002-8439-2616; Rajkai, Kálmán László/0000-0003-4095-774X}
}

@article{MTMT:1290843,
	title = {Vegetation of the dolines in Mecsek mountains (South Hungary) in relation to the local plant communities},
	url = {https://m2.mtmt.hu/api/publication/1290843},
	author = {Bátori, Zoltán and Csiky, János and Erdős, László and Morschhauser, Tamás and Török, Péter and Körmöczi, László},
	doi = {10.3986/ac.v38i2-3.125},
	journal-iso = {ACTA CARSOLOGICA},
	journal = {ACTA CARSOLOGICA},
	volume = {38},
	unique-id = {1290843},
	issn = {0583-6050},
	year = {2009},
	eissn = {1580-2612},
	pages = {237-252},
	orcid-numbers = {Bátori, Zoltán/0000-0001-9915-5309; Erdős, László/0000-0002-6750-0961; Körmöczi, László/0000-0003-1027-3822}
}

@article{MTMT:1222871,
	title = {The effect of the expansion of the clonal grass Calamagrostis epigejos on the species turnover of a semi-arid grassland},
	url = {https://m2.mtmt.hu/api/publication/1222871},
	author = {Somodi, Imelda and Virágh, Klára and Podani, János},
	doi = {10.3170/2008-7-18354},
	journal-iso = {APP VEGE SCI},
	journal = {APPLIED VEGETATION SCIENCE},
	volume = {11},
	unique-id = {1222871},
	issn = {1402-2001},
	abstract = {Question: How does the dominance of Calamagrostis epigejos influence
		species turnover of a grassland?
		Location: Loess grassland at the foothills of Bukk Mountains, Hungary
		(47 degrees 54' N, 20 degrees 35' E).
		Methods: Presence/absence of vascular plants and different performance
		attributes of C. epigejos were recorded in a plot-subplot system
		between 2002 and 2005. Appearance and disappearance rates of grassland
		species were calculated for pairs of consecutive years. 1. Mean
		appearance and disappearance rates were compared in grassland plots
		dominated by C. epigejos and in plots free from this species, based on
		Monte Carlo randomization. 2. Mean appearance rates were assessed for
		categories of C. epigejos performance and their confidence intervals
		were calculated via Monte Carlo randomization. For two performance
		variables (percentage cover and shoot number) analyses were performed
		at two spatial scales.
		Results: 1. C. epigejos-dominated plots differed from unaffected ones
		by significantly lower appearance rates. 2. Change in appearance rates
		was best explained by differences in percentage cover of C. epigejos.
		Coarse-scale C. epigejos performance had a closer correspondence with
		appearance rate change than fine-scale performance. Low level C.
		epigejos performance enhanced appearance rate compared to intact
		stands, while high level performance decreased it, regardless of the
		choice of performance measure.
		Conclusions: C. epigejos lowers species number by hindering
		reappearance of species of the original grassland. This is best
		explained by the increased shading effect at the coarse scale. The
		marked non-linear initial enhancement in appearance rate, however, can
		also be taken as an early sign of future species loss.},
	year = {2008},
	eissn = {1654-109X},
	pages = {187-194},
	orcid-numbers = {Podani, János/0000-0002-1452-1486}
}

@article{MTMT:1156464,
	title = {Microclimate of Karstic Dolines},
	url = {https://m2.mtmt.hu/api/publication/1156464},
	author = {Kevei, Ferencné},
	journal-iso = {ACTA CLIMATOL},
	journal = {ACTA CLIMATOLOGICA},
	volume = {32-33},
	unique-id = {1156464},
	issn = {0563-0614},
	year = {1999},
	pages = {19-27},
	orcid-numbers = {Kevei, Ferencné/0000-0002-4143-8506}
}

@article{MTMT:1866092,
	title = {Social behaviour types, the naturalness and relative ecological indicator values of the higher plants in the Hungarian flora},
	url = {https://m2.mtmt.hu/api/publication/1866092},
	author = {Borhidi, Attila},
	journal-iso = {ACTA BOT HUNG},
	journal = {ACTA BOTANICA HUNGARICA},
	volume = {39},
	unique-id = {1866092},
	issn = {0236-6495},
	year = {1995},
	eissn = {1588-2578},
	pages = {97-181}
}