Alienopterix santonicus sp. n., a metallic cockroach from the Late Cretaceous ajkaite amber (Bakony Mts, western Hungary) documents Alienopteridae within the Mesozoic Laurasia

Cockroaches (Blattaria s. str.) were documented from numerous amber localities around the world, representing both extinct and extant families. Alienopteridae is an extinct cockroach family known only from the Cretaceous of Gondwana (Brazil, Botswana, Myanmar amber) and the Cenozoic of North America. Alienopterix santonicus sp. n. from the Late Cretaceous amber of the Ajka Coal Formation (Bakony Mts, western Hungary) extends the rich geographical distribution of the family into Laurasia during the Mesozoic. As a member of the presumably pollinator cohort Alienopteridae, this species could have played an important role in the Ajka Coal ecosystem during the Santonian. The microrectangular structures of the forewing suggest that the new species likely possessed a metallic colouration already known from the group. Combined with the disruptive body pattern this could have served as an advanced camouflage. The microrectangular structures of the forewing were compared to integument microstructures of extant insects with metallic colouration. Various arthropod taxa are already known from ajkaite, and the new discovery further emphasizes the importance of this amber.