Supplementary data on the Middle Miocene (Badenian) fish assemblage of Nyirad (Hungary): revision and new results on faunal composition and paleoenvironment

In 2016, SzABO & Kocsis reported a diverse Badenian shark and ray fauna from the Nyirad pit (Hungary). Since then a few chondrichthyan taxa have been eliminated from the faunal list, and additional taxa (both chondrichthyans and osteichthyans) have been unearthed at the locality. Some earlier collected chondrichthyan taxa have been revised (e.g., Araloselachus cuspidatus), while further new chondrichthyan taxa (e.g., Alopias exigua, Iago angustidens and Rhynchobatus sp.) occurred in the newly collected material. The recovered osteichthyan remains have been also examined in detail and the material includes Crenidens sp., Dentex sp., Diplodus sp., Pagrus sp., Sparus sp., Labrodon sp., Tri gono don jugleri, Acanthurus sp., Sphyraena sp., an indeterminate trichiurid, and an indeterminate tetraodontiform. The newly assessed fish fauna provides further information for the Badenian paleoenvironment of the Pusztamiske Formation in the Nyirad depositional area.