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Thanks to Mark Shepard and Shirley Nichols and two anonymous reviewers for their constructive comments on the many drafts of this manuscript.\nExport Date: 5 November 2019 \n Correspondence Address: Lucci, G.M.; AgResearch Ruakura Research Centre, Private Bag 3123, New Zealand; email: Gina.Lucci@agresearch.co.nz\nFunding Agency and Grant Number: MBIE [C10X1117]; DairyNZ [C10X1117]; FonterraFonterra [C10X1117]; Beef + Lamb New Zealand [C10X1117]\n Funding text: This work was conducted through the Pastoral 21 Environment Program (C10X1117), jointly funded by MBIE, DairyNZ, Fonterra and Beef + Lamb New Zealand. 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