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Several aspects of this optimization problem have been studied (e.g. improving Medium Access Control (MAC) protocols, routing, energy management) mostly separately, although there is a strong inter-connection between them. In this paper an Artificial Intelligence (AI) based framework is presented to address this problem. Mixed-Observability Markov Decision Processes (MOMDPs) are used to effectively model multiple aspects of WSNs in stochastic environments including MAC in data link layer, routing in network layer, data aggregation, power management, etc. MOMDPs distinguish between full and partial observability, hence they are more efficient than other similar AI methods. The proposed framework provides global optimization of user-defined performance metrics, e.g. minimization of time delay, energy consumption and data inaccuracy. Near-optimal joint network policies are obtained via offline approximation of optimal MOMDP solutions and they are distributed among the individual nodes. Resulting node-policies place effectively no additional computational overhead on nodes in runtime. Experiments evaluate the framework by demonstrating near-optimal solutions for a small-scale WSN in detail in case of given tradeoff criteria. 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