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We dedicate this paper to the memory of our dear friend, the late Matteo Griggio. We are grateful to Jan Hottentot, Wolfgang Vogl, and Hans Winkler for providing us with fishes, and to Roland Sasse, Martina Krakhofer, and Eva Strasser for help with the maintenance of the breeding line. We acknowledge the intellectual and financial support provided by Dustin Penn. 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